Overview
Geology and climate have shaped the development of life tremendously. This has occurred in the form of processes such as the oxygenation of the atmosphere, mass extinctions, tectonic drift, and disasters such as floods and volcanic eruptions. Life, particularly bacteria, has also been able to impact the geological makeup of the planet through metabolic processes.
Overview
The history of life and evolution has been characterized by several key events. These events can be grouped as new hierarchal levels of selection coming into play, as biological units coming together in symbiosis and specialization, or in a number of other ways. Other important events are situations of conflict resolution or information transmission, from the genetic to the cultural level.
Overview
We can use methods of genetic analysis to connect phylogenic information to geographical histories. Human migration has left genetic traces on every continent, and allows us to trace our roots back to Africa. Molecular genetic methods allow us to determine whether or not trait states were ancestral, which can have profound implications for fundamental biological ideas.
Overview
The Tree of Life must be discovered through rigorous analysis. Genetic information is crucial because appearances can be deceiving, and species that look similar can prove to be genetically very dissimilar and not share recent common ancestors. Two criteria, used to determine what the “correct” Tree is, are simplicity and whether the tree maximizes the probability of observing what we actually see.
Overview
Speciation is the process through which species diverge from each other and/or from a common ancestor. There are several definitions of species, most of which focus on reproductive isolation and/or phylogenetic similarities. This can cause some controversy. Speciation can result from geographical separation or ecological specialization. There are stages of speciation in which organisms cluster first into distinct populations before finally becoming different species.
Resources
You may view the Yale Galapagos site at: http://cmi2.yale.edu/galapagos_public
Overview
Sexual selection is a component of natural selection in which mating success is traded for survival. Natural selection is not necessarily survival of the fittest, but reproduction of the fittest. Sexual dimorphism is a product of sexual selection. In intersexual selection, a sex chooses a mate. In intrasexual selection, individuals of one sex compete among themselves for access to mates. Often honest, costly signals are used to help the sex that chooses make decisions.
Overview
Sex allocation is an organism’s decision on how much of its reproductive investment should be distributed to male and female functions and/or offspring. Under most conditions, the optimal ratio is 50:50, but that can change under certain circumstances. Sex allocation determines what sexes sequential hermaphrodites should be at each part of their life as well as how simultaneous hermaphrodites should behave. Some species have more control over the sexes of their offspring than others, and adjust the sex ratios of their offspring depending on the environment and conditions.
Overview
Life history covers three main classes of traits in organisms: age and size at maturity, number and size of offspring, and lifespan and reproductive investment. Organisms must make tradeoffs among these traits that typically cause them to come to evolutionary equilibrium at intermediate values. Life history traits are evolutionary solutions to the ecological problems of the risk of mortality and the acquisition of food, and they are expressed in reaction norms that determine the particular traits that an organism will exhibit when its genes encounter a specific environment during development.
Resources
You may view the Yale Galapagos site at: http://cmi2.yale.edu/galapagos_public
Overview
Genomic conflict arises when the interests of various genomic elements, such as chromosomes and cytoplasmic organelles, are not aligned. These conflicts arise in two situations: either when one unit is contained within another, as a mitochondrion is contained within a cell, or when inheritance is asymmetrical. Genomic conflict can thus occur within a cell, within an organism, or between two organisms, such as a mother and developing fetus. There have been several steps taken to avoid these conflicts in sexual species, including the fairness of meiosis and the uniparental inheritance of cytoplasmic genomes.
Overview
There are several explanations for the evolution of sex and its continued prevalence. One is facilitating the spread of helpful mutations while hastening the removal of harmful ones. Another is expediting resistance against pathogens. Sex does have several costs compared to asex, such as only giving half your genome to offspring, having to find mates, and the risk of predation and STDs. Overall, the benefits outweigh the costs and sex has a firm hold on the majority of the recent branches of the tree of life.
